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from the underlying study[0]:

Spatial patterns of shift

We observed a prominent westward and poleward shift in abundance for most tree species in the eastern United States during the last 30 years (Fig. 2). Of the 86 species studied, 73% shifted their abundance centers westward, of which 65% were statistically significant (P < 0.05), whereas 62% shifted poleward, of which 55% were significant (table S1A). The median longitudinal rate for species that shifted westward (15.4 km per decade) was 40% larger than the median latitudinal rate for species that shifted poleward (11.0 km per decade). The primary direction of species abundance shift was northwest (37% of the species), and the least common direction was southeast (2%).

[0] http://advances.sciencemag.org/content/3/5/e1603055

[edit]: formatting