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From the abstract, this is an unexpected asymmetry:

...we find significant information flows from bot-like accounts to non-bot accounts with behaviour differing between sides. Pro-Russian non-bot accounts are most influential overall, with information flows to a variety of other account groups. No significant outward flows exist from pro-Ukrainian non-bot accounts, with significant flows from pro-Ukrainian bot accounts into pro-Ukrainian non-bot accounts...

Their results don't reflect my experience online, though. I observe heavy activity from pro-Ukrainian accounts that are clearly not bots.