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For me I knew twitter had gone mainstream when I was reminded that the clocks in the UK had gone back (to winter time) by a tweet from Stephen Fry!
This reads like a submarine PR-post. At least Twitter is trying to woo investors and users alike with WSJ articles.
Twitter will be mainstream when everyone's mom and dad doesn't say "huh?" when you say "I Twittered" or "tweeted" something.
twitter is on CNN everyday - if that's not mainstream I don't know what is