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I hit a paywall before this article gave me any idea what actually happened. Fortunately, WSJ also made a blog post about the subject: http://blogs.wsj.com/digits/2012/02/16/how-google-tracked-sa...
If you get to the end of the post, it notes that two Google engineers were actually responsible for patching the loophole in webkit that allows storing third-party cookies after a form submission:

http://trac.webkit.org/changeset/92142

But if you try to read the previous 99% of the blog post you'll see how some shady advertising companies perpetually try to game the system without any user consent.