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I like how this post is broken down into specific categories. The most important tools are under "Analyzing and Improving". Obviously I use google analytics but haven't used Visual Website Optimizer. Is it like Optimizely?
things like crazyegg are good too. you can just use the 30 day trial to test it out.
For monitoring the article mentions Pingdom. I have found New Relic's server monitoring [1] to be quite nice as well.

The nice thing is how it integrates with other New Relic monitoring for your app and that it actually is free.

[1] http://newrelic.com/server-monitoring

Whilst technically true, “at the lowest level”, I find it hard not to take with a pinch of salt any article that describes Heroku as a “free service”.